Shibuya, Sonomi, Keizi Kiritani, és Kenji Fukuda. „Life History Traits of Five Ground Beetle (Coleoptera: Carabidae) Species Common in Honshu Island, Japan”. Acta Zoologica Academiae Scientiarum Hungaricae 66, no. Suppl. (december 28, 2020): 185–198. Elérés november 6, 2026. https://s-web3.mtak.hu/index.php/actazool/article/view/4999.